Fievel is a young Russian mouse separated from his parents on the way to America, a land they think is without cats. When he arrives alone in the New World, he keeps up hope, searching for his family, making new friends, and running and dodging the cats he thought he'd be rid off.
老鼠一家在美国安顿下来后不久,发现他们仍然面临着猫的威胁。这让他们迫切想尝试前往西部的新家园,因为那里承诺着老鼠和猫能和平共处。可惜,作出这番承诺的是个油嘴滑舌的骗子,名叫猫拉·沃尔,他实则包藏着险恶的阴谋。对此一无所知的老鼠一家踏上了西行之旅,而他们真正的猫朋友老虎也紧随其后,决心追随朝同一方向离开的女友。
In the historic melting pot of 19th century New York City, Fievel and the Mousekewitz family are struggling to make their American dream come true. But when a mysterious treasure map leads Fievel and his friends to a secret world of Native American mice living beneath the city, he finds something even more precious than gold and jewels!